Pomona College in Claremont, California and, in 1960 ; Ph.D. from the University of Hawaii ;
Australia
professor of botany at the University of Malaya in Kuala Lumpur from 1965 to 1984 ; Botany Department Chair of the Philadelphia Academy of Natural Sciences ;
Further Place of Activity: China, United States of America, Guam, Marshall Islands, Federated States of Micronesia, Madagascar, Malaysia, Mauritius, Papua New Guinea, the Philippines, and Switzerland.
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